Buried Treasure – Mike Taylor

So kicking off the first in the series of favourite / underappreciated papers is Mike Taylor of SV-POW. Here’s his thoughts on one of his own works:

The paper I look on most fondly is Taylor and Wedel (2013) on “Why sauropods had long necks; and why giraffes have short necks”. I like the snarky title, of course — when I give talks about this subject, I just use the second half — and the subject matter is dear to my heart. But it’s how this paper came together that makes me love it the most.

It started out on a car journey in 2008. All three Wedels were staying with us that summer, as Vicki had a leprosy conference in Bradford. Matt and I visited several museums while they were around. I think it was as we were driving back from Oxford that we started listing the ways that sauropod necks didn’t make mechanical sense to us. Since I was the one driving, Matt took out his notepad and started making lists. “What the hell is going on?”, we asked — and so the embryonic project was dubbed WTH, for “what the hell”.

More than any of our other papers, this one went through really significant revisions. The earliest “complete” version was rather formless: it contained a lot of good stuff, but there was no structure to it. We revised it into an unconventional form with three main sections: “Facts”, “Interpretation” and “Speculation”. At this point, the title was still “What the hell is wrong with you? Mechanical design flaws in the necks of sauropod dinosaurs”.

This was also the basic shape of the version we finally submitted to a journal, though by then it had the more sober (and boring) title “Vertebral morphology and the evolution of long necks in sauropod dinosaurs”. We had a very bad review experience at that journal, which I won’t go over here; but suffice to say that the result was that, having thoroughly reworked it into a form resembling the one we know today, we sent it to a different journal rather than back to the first one. We were bullish about this submission, and pleased to think we were giving a good paper to a journal that could probably use it. So we were rather shocked to find it rejected with reviews that we couldn’t sympathise with — especially one that said “The manuscript reads as a long “story” instead of a scientific manuscript”, which we feel is praise though it was intended as criticism.

We made some revisions in response to those reviews, but by the time we’d done that PeerJ was on the horizon so we sent it there — and after very quick and genuinely helpful reviews, it was published as part of that journal’s first batch: https://peerj.com/articles/36/

We’re really happy with the “story-like” final form of the paper. Our goal was to make something that was not only informative but also fun to read. I hope the progression of the argument makes sense — Introduction, Long Necks in Different Taxa (finishing with sauropods), Factors Enabling Long Necks, Architecture of Sauropod Necks — and that readers always have a solid sense of where they are in the progressing argument. We’re also really happy with the illustrations in this paper: PeerJ, being an online-only open-access journal, imposes no limits, so this is a lavishly illustrated paper with some comparative illustrations (Figs 1, 3 and 7 particularly) that we’re really proud of: https://peerj.com/articles/36/#fig-3

Finally, I won’t deny it’s satisfying that a paper which was (wrongly, we feel) rejected by two palaeo journals has gone on to be viewed 23,000 times by 17,000 different visitors, and has been downloaded 3,000 times. We very much hoped that that paper would reach a non-specialist audience as well as other researchers, and those numbers suggest that’s happening.


Finally, Mike has a pick for an underappreciated paper by someone else is:

Hokkanen, J. E. I. 1986. The size of the largest land animal. Journal of Theoretical Biology 188: 491-499.

New series – Buried Treasure

The Musings has been too quiet of late what with mad work commitments, and my ongoing responsibilities for blogging etc. elsewhere means I have too little time. The old days of a post nearly every day are, I suspect, never coming back but I do want to keep producing material on here. Happily I have a cunning plan (insert your own favourite Blackadder response here) and more happily still, a number of colleagues could be persuaded to write something for me that I can put up here.

Anyone who has read or written a fair amount of scientific papers will know that there are lots of hidden gems out there. Yes, there are tons of celebrated great papers, and tons that all but deserve to be overlooked, but it’s also true that there are many great papers, or even important bits of papers that are glossed over, or simply never spotted. There’s numerous examples of major discoveries turning out to have been already found or worked out years or decades before and even in the modern digital age, people cannot find, let alone read, everything. Important bits of papers, or whole manuscripts will fall by the wayside and key points missed or underappreciated.

With this in mind comes the new series – Buried Treasure (and thanks to Paul Barrett for coming up with the name) where authors talk about papers of theirs or bits of papers which deserve a second (or even a first) reading. Obviously academics are sensitive about their paper and do get annoyed when things are missed or bypassed, so while this isn’t supposed to be a place for axe grinding, (or tooth grinding) it does hopefully provide a platform for people to showcase their work and talk about how papers came about and why they think something is important and might benefit people to revisit it.

The whole thing is supposed to be a bit of fun and rather free from constraints, so people have already suggested they might write about papers that are not their own, but simply one they think needs some more recognition, or just want to write about a paper that has a strong significance for them, or they simply enjoyed writing. Hopefully it’ll be interesting and readers will discover (or rediscover) some nice ideas and see how others look at their own works.

I’ll kick this off with a first entry tomorrow and then it will build up as posts come in, so it is likely to be fairly irregular and I have no idea how long this will run. However, I do already have a small set ready to go, so it won’t die immediately at least and with luck there will be quite a few to come.


Two million (and 3500)

It was never my intention for the Musings to fall quite this silent but between commitments for the book (still available in many bookshops, online, as an e-book and audio-book), the ongoing Guardian blog and in particular my teaching, I’ve rather run out of time to write posts. And let’s be honest, even this one is just a holding pattern post and is mostly just self-congratulatory. Even though I’ve all but stopped posting here, the huge back catalogue of posts on here (over 1000) between them still clock up hits at a decent rate and so just this week the Musings hit 2 000 000 total views. Whoo, go me etc.

However, all is not lost for fans of Dave-based web content as I do at least still tweet quite often and happily this week also saw me hit 3500 followers. If you want to join then you can follow me as @Dave_Hone and currently there’s a huge stream of tweets on my recent trip to the AMNH in New York.

I really do intend to post a bit more on here again in the future and new year should see me with a bit more time and also some papers coming out which will provide something to discuss. I also managed to find time to get to the Bronx Zoo while I was in New York and hope to get a review up of this as it’s a place I had not visited before. In the meantime, thanks to those who still use this site, and I hope it will continue to be a valuable resource for a long time to come, even if my output remains at a fairly diminished level. Till next time, bye.

The Tyrannosaur Chronicles is here!

Well it’s been coming of course but today sees the publication of my first book. I’ve always wanted to write one and now it’s done and I can (sort of) relax. There’s lots of PR stuff ahead and the official book launch tomorrow, but there’s not much to do now except let it go free and hope that most people enjoy it.

I’ve been writing about dinosaurs and palaeo one way or another for nearly 10 years now between various blogs and ventures as well as the odd review paper and book chapter that are for more of a general audience than a typical paper, but this is obviously a much bigger and rather different undertaking. It’s also rather different in that I was writing for something of a different audience (certainly compared to here where I generally assume readers know at least a little anatomy, what a phylogeny is, what the main time periods were etc.) and over a long book you want to introduce quite a few topics and aspects of not just tyrannosaurs, but also their contemporaries and major issues like behaviour, anatomy, local environments, extinction and more. It turned out to be a lot to cover and while trying to keep it interesting for the reader.

Hopefully, I’ve managed that but it is nervy letting this out into the wider world with little control over it. That may sound odd given how much I’ve written online, but with a blog (either here, on Pterosaur.net or on the Guardian) you have a fair idea of who your audience is likely to be, and people will soon leave if they don’t like it. Getting someone to pick up and be immediately drawn to, and then stick with, a whole tome is rather different so obviously I am nervous and curious as to how it goes from here.

The book is very much in the popular science mould and so while I would hope even some academics and researchers would get something from it and enjoy it, really it is aimed squarely at the general public and those with little or no knowledge of dinosaurs or paleontology and even biology in general. As a result, despite the fact that the book is around 85 000 words long, it really doesn’t delve into the tiny details of but tries to cover a broad spectrum of tyrannosaur origins, evolution and their biology. Given my interests there’s quite a lot on ecology and behaviour and there’s a few bits of informed speculation or suggestions that I hope are novel and interesting, but also clearly flagged as such.

It was a huge effort to write all of this while keeping up with a full time academic job and try and keep my other blogs ticking over, and it was also important to try and update things. The last few years have seen a near endless stream of new tyrannosaurs being named and some parts of the book I changed a half dozen times to reflect the addition of new species, and with the book going to print in February, it’s inevitably already out of date thanks to the most recent addition to the ranks of this clade, despite my efforts. Still, I have tried to make this a modern take on tyrannosaurs and I hope I have managed to overcome a few of the more persistent anachronisms and misconceptions about these animals. Anyway, enough of the (brilliant) text and its (brilliant) author, and time to talk about some other aspects of the book and to give a minimal amount of credit to other people.

The book is illustrated by Scott Hartman and there’s around a dozen figures of his scattered through the book, with lots of skeletals (especially of tyrannosaurs, but also various other dinosaurs too) and other little bits, a number of which were done especially for the book, but will be popping up on his website if they haven’t already. I’m obviously especially grateful to Scott for finding the time to do these and putting so much time and effort into them, the book benefits enormously from it.

There is also a colour section in the middle with numerous photos of various specimens and some reconstructions. Plenty of these have been in print in various places before but there are some novel shots and views of various things and I’ve been blessed with the generous assistance of numerous colleagues and friends who have sent in pictures and allowed me to use them. While I’m on the subject therefore I must thank Peter Falkingham, Jordan Mallon, Larry Witmer, Xu Xing, Lu Junchang and Phil Currie for providing various images and also the Royal Tyrrell, LACM, IVPP, Hayashibara, Mongolia Palaeontological, Royal Sasketchewan, Carnegie and New Mexico Museums, and also Don Brinkman, Mark Loewen and Matt Lamanna for helping me negotiate to get a couple of the images. Finally I must also thank Darren Tanke and Chisaka Sakata for the photos of me that are on the covers of the paper- and hardbacks respectively.

Finally with regard to the text I had a series of editors and assistants at Bloomsbury though most especially I want to thank Jim Martin for commissioning the damned thing in the first place and also in particular for supporting my campaign for the colour scheme of the cover. Several friends of mine including Marc Vincent (yes, that one) read through an early draft for me and provided useful feedback and special mention goes to Tom Holtz for reading through it looking for errors (and mercifully he found only one, so I’m happy to blame him for any others that slipped through). A whole host of other friends, collaborators, coauthors and colleagues are thanked in the acknowledgements for sharing their knowledge of tyrannosaurs with me over the years and I hope this book helps do justice to these amazing animals.

Well, the book is out now (actually I’ve had reports of it being on sale since Monday) and while I’ve always wanted to say it’s available in all good bookshops actually I have no idea. It is available online (including direct from the publishers Bloomsbury) and it’s in at least a few physical places. I know it’s available in hardback (paperback coming next year) and e-book versions and there’s an audio version coming via Audible, and hopefully a few translations too. The US have to wait till early June, but not long for you to wait and in the meantime you can enjoy me talking about the book here. Hopefully many people will find it one way or another (such as in charity shops for £2 in a few weeks) but more importantly I do hope people enjoy it. Happy reading.

The Tyrannosaur Chronicles actually exists!

So the official publication date is drawing near (21st of April) of my first book and I actually have a physical copy in my hands! Oooh! It’s got nice pictures and photos and words and everything!

Obviously I’m very pleased but I am also rather nervous about the whole thing – people will be paying actual real money and I really don’t want to let them down. I know you can’t please everyone and even the greatest books will not appeal to every person that picks up and reads even a few pages but despite the years of blogging and outreach stuff this is a new style and form and it’s rather more global in spread than even online media. So, lots of nerves my end.

However, anyone who does buy it and hurls it across the room a few hours later in frustration may at least be mollified by having paid 30% below the cover price thanks to a discount being offered by the publishers. If you order direct from the publishers Bloomsbury before May 31st and enter the promo code ‘DINOSAUR’ at the checkout, it should be reduced. (This has only just been set-up, so do leave a comment if this doesn’t work, or indeed if it does to let me know it’s working!).

Finally, if you are in and around London there is a small formal book launch on the 22nd of April. Tickets are free (but you need to reserve them here in advance). It won’t be long or special, I’ll talk about the book for a bit, answer some questions and sign any copies going (available for sale there, and also at a hefty discount).

Hope to see some regulars there and I do hope you enjoy the book.




Dinosaurs Monster Families


Even people living in London may not know the Horniman Museum which sits in south east London, just a few miles from the famous Crystal Palace dinosaurs. The Horniman is a small museum with an excelletn and old-fashioned natural history section full of bones and taxidermied material but with some great illustrations of development, variation and evolution. There’s a section on human cultures and especially tribal artefacts, a small aquarium in the basement and  a petting zoo and gardens. It’s well worth a visit anytime, but they also regularly have special exhibitions and right now it is the above titled one on dinosaur eggs, nests and babies.



The exhibition is not large but it is excellent. I’ve only included a few snapshots here but hopefully it’s clear that there’s some wonderful specimens (almost all casts, but very few are of specimens or even species I have seen before and none will be well known in the UK), with interesting mounts, excellently presented information and lots of detail. There are some looped videos of researchers talking about major discoveries like the brooding oviraptorosaurs and also lots of top Luis Rey artwork. Luis was actually integral to the origin of this traveling exhibit (it’s also been in Spain and Italy but I don’t know where it’s headed next) and hence the liberal splashing of his works.



Given the theme it’s perhaps no surprise that most of the material is based on Mongolian and northern Chinese specimens – Protoceratops and oviraptorosaurs feature heavily as does Tarbosaurus and innumerable eggs and nests. Again though, while this might in one respect be a bit same-y, you’d have to pay close attention to notice and it’s not played as a central point, merely that so much accessible material is from there so it features. Still there’s stuff from Argentina and North America and lots of key sites and specimens get a mention.


In a nice touch, the last case is a collection of modern specimens from the Horniman’s own collections showing off various bird and their eggs and some other goodies. There’s also a very special ‘guest’ that is quite remarkable to see but I won’t spoil the surprise for anyone going.

The museum also has an excellent record of using these temporary exhibits to carry out additional activities and outreach events, bringing in artists and experts to talk about them to various groups and creating extra activities and presentations. Somewhat inevitably therefore I got roped into this and in the opening week look along a gang of students and colleagues to talk dinosaurs and their biology and evolution and I’m back again in a couple of weeks for another talk.


Overall this is a superb little exhibit, there’s a lot to see, it’s well laid out and there’s some interesting and exciting specimens. It’s well labeled and there’s a lot of information to potentially digest and I can highly recommend it.

What is an adult dinosaur?

Back in early 2015 I took a trip to LA, primarily to catch up with Mike Habib and look at some pterosaur and tyrannosaur material there, but I also took some time to see Andy Farke and Matt Wedel up in Claremont. We chatted about various ideas for things we could collaborate on and threw around a few ideas. Andy suggested something on ontogeny and this soon led to the issue of diagnosing life stages for dinosaurs – something that had been an issue for our Protoceratops paper – and within a few weeks I’d actually had an invitation to submit a review to Biology Letters, and so a plan was hatched.

That paper is now out and in it we look at the vexed issues of what are adult / subadult / juvenile / hatchling etc. dinosaurs. This is of course really quite fundamental to huge amounts of research, if it’s not clear how old an animal is, then issues like taxonomy, systematics and their position in an ecosystem are going to be hard to sort out. Comparing across specimens or species will also have their issues. None of this is a major surprise and yet looking though the literature it’s clear that although people recognise this, they don’t necessarily actually define the nature of the animals they are working on. Things are called ‘adult’ or ‘subadult’ without a definition, specific diagnosis or reference to papers or alternatively they do provide some kind of definition and reason for the assignment but it’s different from all the others out there. It doesn’t take long to find a bewildering and ever changing list of definitions, none of which can be aligned or compared easily between specimens or species.

There’s clearly nothing wrong in principal with diagnosing an animal by different means but not all specimens can be accessed in the same way or preserve things you want to look at. So something that can help bring them into alignment should help everyone. This is a key part of the paper as we try to come up with something close to a universal definition that should apply as widely as possible. We make it very clear that this should be only a starting point and that whatever works for people is fine, but that hopefully it helps, and even if people utterly ignore these definitions, in general we need to be much more careful about actually putting definitions into papers, even for things that are ‘obviously’ adults or juveniles.

Although short, we do cover a lot of ground in the paper and I hope there’s things in there that will resonate and be familiar and useful to many people (and of course lots of the points apply to other extinct clades too). There’s obviously a lot more to come here and more nuance and details than we could easily include but it’s one of he most contentious and important issues around at the moment and I really hope we have contributed meaningfully to it.

The paper currently seems to be available freely online and can be downloaded here.

Edit: here’s a bonus – Mat Wedel’s sauropod-centered take on the paper

Hone, D.W.E., Farke, A.A., & Wedel, M.J. 2016. Ontogeny and the fossil record: what if anything is an adult dinosaur? Biology Letters


Why Jurassic Park III is objectively* the best of the franchise

Every film in the Jurassic Park / World franchise has plenty of problems, but it is actually quite simple to work out which is the best of the four films to date.

Do you know what I want to see in a dinosaur film like Jurassic Park? Dinosaurs.

Do you know what I don’t want to see in a dinosaur film? Annoying children.

So, which of the films has the the most amount of dinosaur footage (absolute, and especially relative to run time) and the least amount of annoying children? Yes, Jurassic Park III is in fact quite clearly the best film to date. Simple. Case closed.

* For a given value of ‘objectively’

Guest Post: Producing Protoceratops art

The little ceratopsian Protoceratops (and indeed art on Protoceratops) has been a big thing for me in recent years as I’ve been lucky enough to work on some very special specimens and have them illustrated in life.  As is so often the case though, one new specimen begets some new opportunities and today sees the publication of a new paper on the ongoing issue of sexual selection and social dominance signals using some of these specimens in the dataset. The paper is freely available online here and I’ve also written about it here, but the paper also contains some lovely new palaeoart of signaling dinosaurs by Rebecca Gelernter who has kindly agreed to talk about her work here.


When I plan a piece of paleoart, I try to make the animal I’m restoring as complete as possible. I want to make it look like a real, tangible creature with adaptations that make sense for its life history. I particularly enjoy showing behavior, which made this a really intriguing project to work on.

First off, I had to figure out what my Protoceratops should look like. Anatomically, this was pretty straightforward, thanks to the wealth of fossil photos, papers, and books Dave had on hand. Factor in his enthusiastic feedback and that’s all the background you could ever need. At Dave’s request, I was depicting the animals without any filaments or other non-scale integument, so after familiarizing myself with the fine points of ceratopsian feet and beaks, all that remained was to design the color scheme.

Proto Sketches

I decided that the facial markings should be only part of the body with elaborate markings, as the frill and jugal bosses were proposed display structures. When designing markings for extinct animals, I like to thumbnail several different possibilities based closely on living creatures and remix them into something new. For Protoceratops, I mostly looked at antelope facial markings, and the final design features elements of bongo and sable. The jugal bosses are an eye-catching white, and the all-important frill is a splash of those ever-popular display colors, orange and red. I imagine that the animal would flush the frill with blood during an encounter with a potential mate or rival for flashier color. I used a camouflage-friendly beige for the animal’s base color, broken up by a line of darker splotches down each side that become bolder and more regular on the tail, another potential display structure. I used white again on the tip and ventral side of the tail to create a starker contrast, with more orange to draw attention to the ridge formed by the tall neural spines.


Dave asked for the piece to show two adult Protoceratops having a confrontation, while a group of less flashy subadults goes about its business in the background. I selected a pose that showed off the display structures: tail up, frill angled toward the other individual. I angled one adult’s head toward the viewer and one away to show that the display colors are limited to the front – no point wasting resources to color the side of your head that you can’t show off. I wanted the piece to be taller and narrower than your standard portrait orientation, so I raised the point of view above the two main animals and arranged the background players some distance away on another dune. Dave suggested adding the crisscrossing footprints in the staging area to suggest that this type of interaction has happened there before. I placed the animals in a particularly empty bit of desert, with just a few small, scrubby plants in the background.

I’d recently gotten good results from painting over a graphite drawing in Photoshop, so I was eager to try that again. There are different ways of doing this, but the technique I usually use is to set the graphite original to “multiply” and leave that layer on top, painting on a few different layers stacked underneath it. It’s an interesting change from using purely traditional media, and I’m looking forward to trying new things with it.

So there you have it: my process for making (definitely) accurate, (hopefully) interesting paleoart. If you’d like to see more of my work, I’m on all the usual sites under the name Near Bird Studios.

Archosaur Musings 2015 Roundup

For the first time I’m breaking away from the previous annual awards and I’m writing something that is more of a general roundup of the year. I already had found I needed to heavily alter my previous series of awards last year with my changing interests and responsibilities and finding that I’d need to make even more drastic edits this year I though it time to finally shelve the awards and move to a more general summary of the year.

As with last year my blogging has been even more sparse. In part this is down t having less and less time available and also the fact that I have now written close to 2000 pieces between the Musings, the very old (and now apparently no longer online) Dinosaur index on Bristol University’s system, my Guardian blog and various other outlets. That’s on top of the 1000+ questions I’ve answered on Ask A Biologist as well and it all means that I’m somewhat worn down by blogs. Not that I don’t have a desire to continue, but it’s hard not to rehash existing issues and the most popular areas (bird origins, new species) are very well covered and I struggle to bring anything new or find the enthusiasm a lot of the time.

Still, things are continuing. People might have noticed that the Guardian blog in particular has been in hibernation for around 6 months now. It was originally my intention to quit as while I liked it, there were ever increasing pressures to cover the very areas I had least interest in but a solution was stumbled upon – to draw in additional bloggers and expand this from just dinosaurs to all palaeontology. As such there was a call out for people to apply and the editors are close to making a decision on who will be asked to join me and the whole thing should restart in the new year – stay tuned.

My own new year for 2015 saw me taking a trip to LA for a long overdue break, to see the LACM and its collections, visit La Brea and its tar pits and in particular catch up with Mike Habib and try to finish off some papers. Our work on a new and exceptional Rhamphorhynchus held in Canada is now out, as is out collaboration with artist Matt van Rooijen on wingtip curvature and what that means both ecologically and perhaps systematically for pterosaurs.

Sadly for me this summer lacked any meaningful trips – I’ve been out of the field far too long, and I desperately need to get back to China to finish several projects, but the late summer saw a flurry of activity. First off, rising artist Rebecca Gelernter joined me in London for several months to work on a series of projects as part of her scientific illustration degree. Some of her work (both life reconstructions and skeletal work) will be appearing very shortly in a number of papers for me and John Hutchinson has also put her nose to the grindstone for some illustrations too. If you’ve not seen her stuff before, do take a look at her website and she recently joined Twitter too.

Next was an obvious highlight of back to back conferences: Flugsaurier in Portsmouth and SVPCA in Southampton. The former was the latest in the running series of pterosaurs conferences and saw a superb collection of talks as well as the obvious benefit of getting together people from all over the world to talk pterosaurs. Seeing colleagues and experts you may only otherwise rarely or never see makes it an extremely valuable gathering, even if there were no talks and posters. Still, much was exchanged and much got done and a great time was had by most who survived the weather. As is also becoming a pattern, a volume of papers will also be published from this meeting, and well follow the link if you want more.

SVPCA was a bit more cosmopolitan than usual as several pterosaur delegates stayed on for the second meeting (as had been hoped, each meeting encouraged some people to the other when they would not normally attend) but was also an excellent meeting and gathering of vertebrate palaeontologists. There were some format changes (with more to come) but none the worse for it, and for me it is probably the best annual meeting out there and I love it. Long may it continue, though sadly I look set to miss the 2016 meeting owing to being in Canada.

One other thing that needs a mention for 2015 is the Daspletosaurus paper. This started as a crowdfunded platform that took me to Alberta to work on a very chewed-up skull with Darren Tanke. It took a while but the paper was eventually completed and published and I’m very pleased with the final, detailed study. A lot of people contributed their time as well as cold, hard cash and I’m extremely grateful for all the help that allowed me to complete this research.

Looking ahead, I’m working on what are hopefully the final edits on the Tyrannosaur Chronicles that will be my first book, and there’s a paper on sexual selection in dinosaurs now in press that should be out in the next few weeks. There’s a couple of other works in submission and I’m contributing to the Flugsaurier volume too, so fingers crossed that I’ll have a couple more pieces out next year. That pretty much wraps it up for now. This blog will continue sporadically I’m sure so keep an eye out for new posts.

Flugsaurier 2015 Volume of Papers announced

I’m putting this up here as I do still get a fair number of visitors and obviously this site is still very pterosaur centric. Pterosaur enthusiasts will know there have been a series of volumes of pterosaur reseach stemming from the various conferences in the last decade or so. First and foremost among them has been the Geological Society’s special volume from 2003 that came from the meeting in France. Happily the same venue have agreed to publish the next in the series which is based on the the recent conference in Portsmouth. Potential authors should read on!


Dear Authors,

We have now had confirmation that the Geological Society will be publishing a special volume of papers centred around the 2015 Flugsaurier conference that was recently held in Portsmouth, UK. This volume will be edited by Dave Hone, David Martill and Mark Witton.

The Society have set a deadline of the 31st of January 2016 (a Sunday) for the first submission of manuscripts. This is relatively short notice, but includes the Christmas and New Year periods when traditionally teaching commitments are relatively low. At nearly 3 months away, hopefully this not too onerous a deadline. Formatting and submission instructions can be found here: www.geolsoc.org.uk/sp_authorinfo

Manuscripts will be taken through a full editorial process and subjected to peer review. Manuscripts may be rejected or subjected to multiple rounds of review if their content is not endorsed by referees. Unlike the very successful 2003 Geological Society pterosaur volume, accepted manuscripts will be published online immediately after review and corrections. Volume contents will therefore not be held up by any delays surrounding single manuscripts. A hard copy volume will be printed once all manuscripts are accepted, the deadline for which is 2017.

The volume has capacity for a large number of manuscripts and we encourage submissions. If space does become an issue, we will prioritise content related to material presented at Flugsaurier 2015, and authors who have already announced an intention to submit to the collection.

Please do contact us if you have any questions or queries. We look forwards to reading you research.


Dave, David and Mark



Pterosaur wingtips – not on the straight and narrow

Take a look at almost any illustration of a pterosaur, be it in a research piece or a life reconstruction and the wing finger is generally depicted as being some kind of straight spar. Each of the four wing finger bones is a dead straight element and the leading edge is therefore basically just a line drawn with a ruler). However, take a look at the actual specimens of pterosaurs and it’s actually quite clear that for lots of them, the last (distal) element is often curved, if only a little, but sometimes quite a lot.

This is really obvious in something like Pteranodon for example (and indeed it’s been noted before that this genus has curved distal phalanges) and yet illustrations of this animal, even in the technical literature, will give it a straight distal phalanx. I’d noted for a while that actually there were quite a few pterosaurs with curved phalanges in particular having looked at Bellubrunnus and its bizarre forward swept wingtips. I’d realised that even the posterior curve might actually have some major flight implications – the shape and position of the very distal part of the wing can have a big impact on vortex shedding and other issues even in static glides and anything like a twist or elevation to the tip can make a huge difference to how it performs.

Knowing this would be an issue and working out what it would be and why are two very different areas and I know enough mechanics for the first and not enough to even begin to think about the second. Enter, somewhat inevitably, Mike Habib and he started looking at this issue and working towards what such a curve would mean both in Bellubrunnus but also those pterosaurs with posterior curves on the distal phalanges. We still needed a good dataset and some actual numbers though and so while I trawled the literature and my photographic archives for examples, any I found I passed onto Matt Van Rooijen who had volunteered to produce both the figures for the paper but also do the detailed digital measuring of the curvature of the phalanges.

The resultant paper is rather light on in depth analysis and numbers because there are potentially some severe issues of taphonomy that can distort the apparent curvature of these bones (in particular reducing a curved bone to look straight) but given the strong consistency of at least some results, there do appear to be some major and genuine signals in the data. There’s some fair consistency within and between clades therefore (and to a degree within and between species of a single genus) so despite the taphonomic issue, it’s perhaps not too bad (though still very hard to estimate or account for).

A number of specimens of multiple genera show that scaphognathines and tapejarids have relatively strong curvature to the distal phalanges and so to do various pteranodontids. In other words, two groups often considered to be highly terrestrial, and another than is highly pelagic both seem to go more for this curvature and others show lesser or no curvature. This might seem rather odd with the two extremes of flying environment / style coming together in morphology but it actually makes a fair bit of sense.

Curvature in the pteranodontids would potentially correspond to an expanded wingtip which aligns with existing hypotheses of the forward swept wing position of these animals in flight. A curved wingtip can also increase the chord of the wing which would be good for terrestrial-based fliers, and also might help protect the wingtip from damage from impact which could be important for animals flying in cluttered environments.

An additional issue comes in here of compliance, a compliant phalanx could potentially also help reduce injuries from impact with things like twigs or even the ground when taking off. Bat phalanges are highly compliant (i.e. bendy) under loads but eyeballing bat fossils at least, there’s no obvious difference between the bones of the phalanges and other elements of the skeleton that are less compliant, so perhaps at least some pterosaur phalanges were highly compliant. In that case under loading in flight they could be considerably more curved, and those of Bellubrunnus might actually be straight in flight!

Overall then this paper has a bit of something for everyone (hopefully). There is likely to be some kind of taxonomic and systematic signal in the presence of curved wingtips though it would have to be treated with caution as a potential character, but that’s also true of lots of other things too, it should not be overlooked. Second, there really does seem to be an ecological signal there which helps potentially restore the ecological habits and habitats of various taxa. There is very much some aerodynamic ideas in here which can be explore further in terms of wingtip shape, and the implications for thing like chord, stall speeds and how this might relate to wing position in flight. Out hypothesis about compliant bone can potentially be tested with histological sampling and finally this should provide a bit more information for those of the artistic persuasion who like drawing pterosaurs. Enjoy!

Hone, D.W.E., van Rooijen, M.K., & Habib, M.B. 2015. The wingtips of pterosaurs: anatomy, aeronautical function and ecological implications. Palaeogeography, Palaeoclimatology, Palaeoecology, 440: 431-439.

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